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There is limited time for elementary teacher professional learning in science in order to meet the aspirational goals of current reform efforts. In this study, we investigated what and how teachers learn on-the-job to gain insight into modes of support less often included in teacher education design. Specifically, we studied elementary teachers’ participation in a system-level organizational routine: curriculum materials adoption processes(CMAPs).Using a communities of practice framework, we explored teacher learning in a comparative case study of three U.S.school districts’ CMAP routines, observingCMAP committee meetings and interviewing participants about their expeirences. We found that what teachers learned varied across each district’s CMAP. We argue this variation can be traced to two CMAP features:(1) teachers’ use of boundary objects and (2) their boundary spanning roles and structures. Results have implications for the design of educational systems’ organizational routines to more intentionally serve a dual role as a professional learning opportunity.